Thumb-up or Thumbs-up

Thumbs for me.
 
Yep, "thumbs-up" for me, too, since it's not just a thumb being up, as it were, it's a pictorial form of the recognised thumbs-up sign.
 
I don't have a clue as to why it would be a plural and not a singular.
I'd never thought of this before, but yes, weird. You do sometimes see people do it with both thumbs, but I thought the origins of thumbs up or thumbs down was (at least in popular imagination) the old Roman gladiatorial thing, and that was only one hand, I think? But you always hear "He gave him the thumbs down".
 
I think it's plural as the origins of the term seem to be describing the action of more than one person; the thumbs of the crowd at the coloseum or the thumbs of the archers indicating they are ready for battle. Only later did it get associated with a single person and a single thumb.
 

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